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Publications

Tau Induces Taxol to Bind Microtubules Cooperatively

Accepted for publication in Proceedings of the National Academy of Sciences

ABSRTACT: Taxol and tau are two ligands which stabilize the microtubule lattice. Taxol is an anti-mitotic drug that binds b tubulin in the microtubule interior. Tau is a microtubule-associated protein that binds both a and b tubulin on the microtubule exterior. Both taxol and tau reduce microtubule dynamics and promote tubulin polymerization. Tau alone also acts as a buttress to bundle, stiffen, and space microtubules. A structural study recently suggested that taxol and tau may interact by binding to the same site. Here, we report that tau induces taxol to bind microtubules cooperatively dependent on the tau concentration using fluorescence recovery after photobleaching. We develop a model that correctly fits the data in the absence of tau and yields a measure of taxol cooperativity when tau is present.

Mobility of Taxol in Microtubule Bundles

Accepted for publication in Biophysical Journal; Vol 84, June 2003.

ABSTRACT: Mobility of taxol inside microtubules was investigated using fluorescence recovery after photobleaching (FRAP) on flow-aligned bundles. Bundles were made of microtubules with either GMPCPP or GTP at the exchangeable site on the tubulin dimer. Recovery times were sensitive to bundle thickness and packing, indicating that the microtubule wall is permeable to taxol. The density of open binding sites along a microtubule was varied by controlling the concentration of taxol in solution. With > 63% sites occupied, recovery times were independent of taxol concentration and, therefore, inversely proportional to the microscopic dissociation rate, koff. It was found that 10 koffGMPCPP ~ koffGTP, consistent with, but not fully accounting for, the difference in equilibrium constants for taxol on GMPCPP and GTP microtubules. With < 63% sites occupied, recovery times decreased as for both types of microtubules. We conclude that the diffusion of taxol inside the microtubule bundle is hindered by rebinding events when open sites are within ~7 nm of each other.

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Taxol Mobility

Presentations

"Tau Induces Taxol to Bind Microtubules Cooperatively"
APS March Meeting. Montreal, CANADA 10:00am March 23, 2004

"Tau Induces Taxol to Bind Microtubules Cooperatively"
Poster Presentation Biophysical Society Meeting Baltimore, MD. February 17, 2004

"Tau Induces Taxol to Bind Microtubules Cooperatively"
Condensed Matter, Applied Physics Seminar. University of California, Santa Barbara, CA. January 8, 2004

"Diffusion Inside and In Between Microtubules"
Institute for Medicine and Engineering Chalk Talk. University of Pennsylvania, Philadelphia, PA. June 13, 2003

"Using FRAP on Microtubules - recent work update"
Fyl and Fredrickson's group meeting. University of California, Santa Barbara, CA. April 25, 2003

"Macromolecular Diffusion In and Around Microtubules"
Condensed Matter and Applied Physics Seminar. University of California, Santa Barbara, CA. April 24, 2003

"Affects of Tau on Taxol Mobility"
12 minute talk. APS March Meeting, Austin, TX. 8:00am Wednesday, March 5, 2003

"Mobility of Taxol Inside Microtubule Bundles"
Poster Presentation. Biophysical Society Meeting, San Antonio, TX. TBA

"Mobility of Taxol Inside Microtubule Bundles"
Condensed Matter Seminar. University of Pennsylvania, Philadelphia, PA. October 16, 2002

"Random Walks Through Microtubules"
Invited Seminar. Wellesley College Physics Department, Wellesley, MA. October 9, 2002

"Random Walks Through Microtubules: Mobility of Taxol Inside Microtubule Bundles"
Advancement to Candidacy Exam. University of California, Santa Barbara, CA. October 1, 2002

"Mobility of Taxol in Microtubule Bundles"
Poster Presentation. Departments of Molecular, Cellular and Developmental Biology (MCDB) and Biomolecular Sciences and Engineering (BMSE), University of California, Santa Barbara, CA. September 20, 2002

Awards, Fellowships, and Research Opportunities

UCSB Affiliates Barbarba S. Uehling Graduate Dissertation Fellowship, declined
University of California, Santa Barbara 2004

Materials Research Laboratory Graduate Student Travel Grant,
University of California, Santa Barbara 2003

Ferrando-Fithian Fellowship for Outstanding Woman in Physics,
University of California, Santa Barbara 2001-2002

Graduate Assistance in the Areas of National Need (GAANN) Fellowship,
University of California, Santa Barbara 2000-2001

Phyllis Flemming Physics Prize for Outstanding Physics Major,
Wellesley College 2000

Jerome A. Schiff Fellowship for Thesis Work,
Wellesley College. 1999-2000

Massachusetts Spaces Grant Consortium Thesis Fellowship,
Wellesley College. 1999-2000

Hughes Research Travel Grant,
Wellesley College. 1999-2000

Outstanding Student Presentation,
MathFest, Providence, RI. 1999

NASA Reduced Gravity Student Flight Opportunities Program,
"Video of Physics in Reduced Gravity Used for Education" 1998-1999
"Group Investigating Rotating Liquids under Space Conditions just wanna have Fluids Under No-g" 1999-2000